Results/Conclusions: CHAR in grasslands indicates shifts in factors such as fuel availability, fuel moisture, and/or vegetational type. CHAR remains low (<2.9 pieces/cm2/yr) between 25-14 ka at Rutundu and is higher and more variable (0.8-13.5 pieces/cm2/yr) at Challa. However, pollen assemblages indicate that open grassland prevailed near both lakes at this time. These patterns suggest greater and more variable biomass burning at Challa probably because of greater moisture availability and warmer temperatures leading to higher plant productivity. CHAR in both lakes increase in parallel with an increase in regional moisture availability after ~16 ka. The convergence in the CHAR records likely represents an increase in fuel availability as a result of a shift to closed bushed grassland, and thus increased plant biomass, around Rutundu. CHAR reaches its maximum values in both lakes ~10.5 ka and then generally decreases as East Africa grew drier towards the present. The CHAR records also reveal fire-regime shifts coincident with abrupt changes during the last glacial-interglacial transition. For example, CHAR in both lakes decline at 12.9 ka and then rise rapidly at 11.5 ka in concert with fluctuating moisture levels associated with Younger Dryas (YD) aridity. At Challa CHAR fluctuations coincide with centennial-scale events in the GISP2 d18O record, including the Intra-Allerod Cold Period (~13.1 ka) and the Pre-Boreal Oscillation (~11.4 ka). These results suggest a connection between northern hemisphere climate variability and fire regimes in East African grasslands.