Natural communities are often characterized
by species rich, reticulate food webs.
Efforts to estimate the strength of species interactions in such systems
have been hampered by the multitude of direct and indirect interactions their
food webs exhibit and have been limited by an assumption that the strengths of
pairwise species interactions display linear functional forms. Here we present a new method for directly
measuring, on a per capita basis, the nonlinear strength of trophic species
interactions within such food webs. The
approach of this method is observation-based, requiring three pieces of
information obtainable through field surveys and laboratory experiments: (1)
species abundances, (2) predator-prey-specific handling-times, and (3)
predator-specific feeding-surveys that tally the number of predator individuals
observed to be feeding and not feeding on each of the predator's prey species
at any given time. In this talk we
describe the advantages and assumptions of the method, and apply it to data
from a